The scandinavian brown bear research project

A66 Dahle, B., O.-G. Støen, and J. E. Swenson. 2006. Factors influencing home-range size in subadult brown bears. Journal of Mammalogy 87:859-865.

A67 Støen, O.-G., A. Zedrosser, P. Wegge and J. E. Swenson. 2006. Socially induced delayed primiparity in brown bears Ursus arctos. Behavioural Ecology and Sociobiology 61:1-8.

A68 Miquel, C, Bellemain, E., Poillot, C, Bessière, J., Durand, A. and Taberlet, P. 2006. Quality indexes to assess the reliability of genotypes in studies using noninvasive sampling and multiple-tube approach. Molecular Ecology Notes 6:985-988.

A69 Gervasi, V., S. Brunberg, and J. E. Swenson. 2006. An individual approach to measure activity levels: a test on brown bears. Wildlife Society Bulletin 34:1314-1319.

A70 Bellemain, E., Nawaz, M. A., Valentini, A. Swenson, J. E., Taberlet. 2007. Genetic tracking of the brown bear in northern Pakistan and implications for conservation. Biological Conservation 134:537-547.

A71 Zedrosser, A., Bellemain, E., Taberlet, P., and Swenson, J. E. 2007. Genetic estimates of annual reproductive success in male brown bears: the effects of body size, age internal relatedness and population density. Journal of Animal Ecology 76:368-375.

A72 Manel, S, F. Berthoud, E. Bellemain, M. Gaudeul, G. Luikart, J. E. Swenson, L. P. Waits, P. Taberlet and Intrabiodiv Consortium. 2007. A new individual-based spatial approach for identifying genetic discontinuities in natural populations. Molecular Ecology 16:2031-2043.

A73 Nellemann, C., O.-G. Støen, J. Kindberg, J. E. Swenson, I. Vistnes, G. Ericsson, J. Katajisto, B. P. Kaltenborn, J. Martin, and A. Ordiz. 2007. Terrain use by an expanding brown bear population in relation to age, recreational resorts and human settlements. Biological Conservation 138:157-165.

A74 Swenson, J. E., M. Adamič, D. Huber, and S. Stokke. 2007. Brown bear body mass and growth in northern and southern Europe. Oecologia 153:37-47.

A75 Moe, T. F., Kindberg, J., and J. E. Swenson. 2007. The importance of diel behaviour when studying habitat selection; examples from female Scandinavian brown bears (Ursus arctos). Canadian Journal of Zoology 85:518-525.

A76 Swenson, J. E., B. Dahle, H. Busk, O. Opseth, T. Johansen, A. Söderberg, K. Wallin and G. Cederlund. 2007. Predation on moose calves by European brown bears. Journal of Wildlife Management 71:1993-1997.

A77 Nawaz, M. A. 2007. Status of the brown bear in Pakistan. Ursus 18:89-100.

A78 Zedrosser, A., Støen, O.-G., Sæbø, S., and Swenson, J. E. 2007. Should I stay or should I go? Natal dispersal in the brown bear. Animal Behaviour 74:369-376. (Featured article)

A79 Ordiz, A., C. Rodríguez, J. Naves, A. Fernández, D. Huber, P. Kazcensky, A. Mertens, Y. Mertzanis, A. Mustoni, S. Palazón, P. Y. Quenette, G. Rauer, C. Rodríguez, and J. E. Swenson. 2007. Distance-based criteria to identify minimum number of brown bear females with cubs in Europe. Ursus 18 :158-167.

A80 Fredriksson, G. M., Danielsen, L. S., and Swenson, J. E. 2007. Impacts of El Niño related drought and forest fires on sun bear fruit resources in lowland dipterocarp forest of East Borneo. Biodiversity and Conservation 16:1823-1838.

A81 Bischof, R., Swenson, J.E., Fujita, R., Zedrosser, A., and Söderberg, A. 2008. Hunting patterns, the ban on baiting, and harvest demographics of brown bears in Sweden. Journal of Wildlife Management 72:79-88. (Featured article)

A82 Bostedet, G., Ericsson, G. & Kindberg, J. 2008. Contingent values as implicit contracts – estimating minimum legal willingness to pay for conservation of large carnivores in Sweden. Environmental and Resource Economics 39:189-198.

A83 Ericsson, G., Bostedet, G. & Kindberg, J. 2008. Wolves as a symbol for people’s willingness to pay for large carnivore conservation. Society and Natural Resources 21: 21:294-309.

A84 Bischof, R., A. Mysterud, and J. E. Swenson. 2008. Should hunting mortality mimic the patterns of natural mortality? Biology Letters 4: 307-310. (Featured article)

A85 Elfström, M., Swenson, J. E., and Ball, J. P. 2008. Selection of denning habitats by Scandinavian brown bears. Wildlife Biology 14:176-187.

A86 Swenson, J. E. and Haroldson, M. A. 2008. Observations of mixed-age litters in brown bears. Ursus 19: 73-79.

A87 Jombart, T., S. Devillard, A.-B. Dufour and D. Pontier. 2008. Revealing cryptic spatial patterns in genetic variability by a new multivariate method. Heredity 101:92-103.

A88 Nawaz M.A., Swenson J.E. and Zakaria V. 2008. Pragmatic management increases a flagship species, the Himalayan brown bears, in Pakistan’s Deosai National Park. Biological Conservation 141:2230-2241.

A89 Martin, J., C. Calenge, P.-Y. Quenette, D. Allainé. 2008. Importance of movement constraints in habitat selection studies, Ecological Modelling 213:257-262.

A90 Ordiz, A., O.-G. Støen, J. E. Swenson, I.Kojola, and R. Bischof. 2008. Distance-dependent effect of the nearest neighbor: spatiotemporal patterns in brown bear reproduction. Ecology 89:3327-3335.

A91 May, R., van Dijk, J., Wabakken, P., Linnell, J. D. C., Swenson, J. E., Zimmerman, B., Odden, J., Pedersen, H. C., Andersen, R. and Landa, A. 2008. Habitat differentiation within the large-carnivore community of Norway’s multiple-use landscapes. Journal of Applied Ecology 45:1382-1391.

A92 Valentini, A, C. Miquel, M. A. Nawaz, E. Bellemain, E. Coissac, F. Pompanon, L. Gielly, C. Cruaud, G. Nascetti, P. Winker, J. E. Swenson, and P. Taberlet. 2009. New perspectives in diet analysis based on DNA barcoding and parallel pyrosequencing: the trnL approach. Molecular Ecology Resources 9:51-60.

A93 Kindberg, J., G. Ericsson, and J.E. Swenson. 2009. Monitoring rare or elusive large mammals using effort- corrected voluntary observers. Biological Conservation 142:159-165.

A94 Strömquist, A., Å. Fahlman, J. M. Arnemo, and A. Pettersson. 2009. Dental and periodontal health in free-ranging Swedish brown bears (Ursus arctos). Journal of Comparative Pathology 141:170-176.

A95 Bischof, R., J. E. Swenson, N. G. Yoccoz, A. Mysterud, and O. Gimenez. 2009. The magnitude and demographic selectivity of natural and multiple anthropogenic mortality causes in hunted brown bears. Journal of Animal Ecology 78:656-665.

A96 Pagès, M., C. Maudet, E. Bellemain, P. Taberlet, and S. Hughes. 2009. A system for sex determination from degraded DNA: a useful tool for palaeogenetics and conservation genetics of ursids. Conservation Genetics 10:897-907.

A97 Bischof, R., A. Zedrosser, S. Brunberg, and J. E. Swenson. 2009. A note on opportunism and parsimony in data collection. Journal of Wildlife Management 73:1021-1024.

A98 Bischof, R., A. Mysterud, and J. E. Swenson. 2009. With or without equations: What are the dos and don’ts of hunting? Biology Letters 5:213.

A99 Zedrosser, A., B. Dahle, O.-G. Støen, and J. E. Swenson. 2009. The effects of primiparity on reproductive performance in the brown bear. Oecologia 160:847-854.

A100 Kübber-Heiss, A., Zedrosser, A., Rauer, G., Zenker, W., Schmidt, P. and Arnemo, J.M. 2009. Internal hydrocephalus combined with pachygyria in a wild-born brown bear cub. European Journal of Wildlife Research 55:539–542.

A101 Bischof, R and Zedrosser, A. 2009. The educated prey: consequences for exploitation and control. Behavioral Ecology 20:1228–1235.

A102 Elfström, M. and Swenson, J. E. 2009. Effects of sex and age on den site use by Scandinavian brown bears. Ursus 20:85-93.

A103 Ordiz, A., O-G. Støen, L. G. Langebro, S. Brunberg, and J. E. Swenson. 2009. A practical method for measuring wildlife horizontal cover. Ursus 20:109-113.

A104 Nawaz, M. A., J. E. Swenson, and V. Zakaria. 2009. Recovery of brown bears in northern Pakistan. Current Conservation 3:12-13.

A105 Mysterud, A and Bischof, R. 2010. Can compensatory culling offset undesirable evolutionary consequences of trophy hunting? Journal of Animal Ecology 79:148-160.

A106 Fahlman, Å., Pringle, J., Arnemo, J. M., Swenson, J. E., Brunberg, S., and Nyman, G. 2010. Treatment of hypoxemia during anesthesia of brown bears (Ursus arctos). Journal of Zoo and Wildlife Medicine 41:161-164.

A107 Fernández-Gil, A., A. Ordiz and J. Naves. 2010. Are Cantabrian brown bears recovering? Ursus 21:121-124.

A108 Fröbert, O., K. Christensen, Å. Fahlman, S. Brunberg, J. Josefsson, E. Särndahl, J. E. Swenson, J. M. Arnemo. 2010. Platelet function in brown bear (Ursus arctos) compared to man. Thrombosis Journal 8:11.

A109 Martin, J., M. Basille, B. Van Moorter, J. Kindberg, D. Allainé, and J. E. Swenson. 2010. Coping with human disturbance: spatial and temporal tactics of the brown bear (Ursus arctos). Canadian Journal of Zoology 88:875-883.

A110 Karamanlidis, A. A., E. Drosopoulou, M. de Gabriel Hernando, L. Georgiadis, L. Krambokoukis, S. Pllaha, A. Zedrosser and Z. Scouras. 2010. Noninvasive genetic studies of brown bears using power poles. European Journal of Wildlife Research 56:693-702.

A111 Gandolf, A. R., Å. Fahlman, J. M. Arnemo, J. L. Dooley, and R. Hamlin. 2010. Baseline normal values and phylogenitic class of the electrocardiogram of anesthetized free-ranging brown bears (Ursus arctos). Journal of Wildlife Diseases 46:724-730.

A112 Støen, O.-G., W. Neumann, G. Ericsson, J. E. Swenson, H. Dettki, J. Kindberg, and C. Nellemann. 2010. Behavioural response of moose Alces alces and brown bears Ursus arctos to direct helicopter approach by researchers. Wildlife Biology 16: 292-300.

A113 Rosell, F., Jojola, S. M., Ingdal, K., Lassen, B.A., Swenson, J. E., Arnemo, J. M, & Zedrosser, A. 2011. Brown bears possess anal sacs and secretions may code for sex. Journal of Zoology 283:143-152.

A114 Swenson, J. E., P. Taberlet, and E. Bellemain. 2011. Genetics and conservation of European brown bears. Mammal Review 41:87-98.

A115 Fahlman, Å., Arnemo, J. M., Swenson, J. E., Brunberg, S., Pringle, J., and Nyman, G. 2011. Physiologic evaluation of capture and anesthesia with medetomidine-zolazepam-tiletamine in brown bears (Ursus arctos). Journal of Zoo and Wildlife Medicine 42:1-11.

A116 Ordiz, A., O-G. Støen, M. Delibes, and J. E. Swenson. 2011. Predators or prey? Spatio-temporal discrimination of human-derived risk by brown bears. Oecologia 166:59–67.

A117 Vestergaard, P., O.-G. Støen, J. E. Swenson, L. Mosekilde, L. Heickendorff, and O. Fröbert. 2011. Vitamin D status, bone and connective tissue turnover in brown bears (Ursus arctos) during hibernation and the active state. PLOS One 6 (6): e21483.

A118_118 Sahlén, E., O.-G. Støen, and J. E. Swenson. 2011. Brown bear den site concealment in relation to human activity in Sweden. Ursus 22:152-158.

A119 Zedrosser, A., Steyaert, S. M.J.G., Swenson, J. E. and Gossow, H. 2011. Brown bear conservation and the ghost of persecution past. Biological Conservation 144:2163-2170.

A120_120 Kindberg, J., J. E. Swenson, G. Ericsson, E. Bellemain, C. Miquel, P. Taberlet. 2011. Estimating population size and trends of the Swedish brown bear (Ursus arctos) population. Wildlife Biology 17: 114-123.

A121 Karamanlidis, A. A., A. Sanopoulos, L. Georgiadis, A. Zedrosser. 2011. Structural and economic aspects of human – bear conflicts in Greece. Ursus 22:141-151.

A122_122 Steyaert S. M. J. G, Støen O-G, Elfström M, Karlsson J, Van Lammeren R, Bokdam J, Zedrosser, A, Brunberg, S., and Swenson J. E. 2011. Resource selection by sympatric free-ranging dairy cattle brown bears Ursus arctos. Wildlife Biology 17: 389-403.

A123 Fink, T., J. G. Rasmussen, J. Emmersen, L. Pilgaard, Å. Fahlman, S. Brunberg, J. Josefsson, J. M. Arnemo, V. Zachar, J. E. Swenson, and O. Fröbert. 2011. Adipose-derived stem cells from the brown bear (Ursus arctos) spontaneously undergo chondrogenic and osteogenic differentiation in vitro. Stem Cell Research 7:89–95.

A124_123 Bischof, R. and Swenson, J. E. 2012. Combining noninvasive genetic sampling and traditional monitoring to aid management of a trans-border carnivore population. Ecological Applications 22:361-373.

A125 Steyaert, S. M.J.G., A. Endrestøl, K. Hackländer, J. E. Swenson, and A. Zedrosser. 2012. The mating system of the brown bear Ursus arctos. Mammalian Review 42:12-34.

A126 Gervasi, V., E. B. Nilsen, H. Sand, M. Panzacchi, G. R. Rauset, H. C. Pedersen, J. Kindberg, P. Wabakken, B. Zimmermann, J. Odden, O. Liberg, J. E. Swenson, and J. D. C. Linnell. 2012. Predicting the potential demographic impact of predators on their prey: a comparative analysis of two carnivore–ungulate systems in Scandinavia. Journal of Animal Ecology 81:443-454.

A127 Arinell, K., B. Sahdo, A. L. Evans, J. M. Arnemo, U. Baandrup, and O. Fröbert. 2012. Brown bears (Ursus arctos) seem resistant to atherosclerosis despite highly elevated plasma lipids during hibernation and active state. Clinical and Translational Science 5:269-272.

A128_128 Moen, G. K., O.-G. Støen, V. Sahlén, and J. E. Swenson. 2012. Behaviour of solitary adult Scandinavian brown bears (Ursus arctos) when approached by humans on foot. PLOS One 7(2): e31699.

A129_129 Gonzalez, O., A. Zedrosser, F. Pelletier, J. E. Swenson, and M. Festa-Bianchet. 2012. Litter reductions reveal a trade-off between offspring size and number in brown bears. Behavioral Ecology and Sociobiology 66:1025-1032.

A130 Fahlman, Å., N. Caulkett, J. M. Arnemo, P. Neuhaus, and K. E. Ruckstuhl. 2012. Efficacy of a portable oxygen concentrator with pulsed delivery for treatment of hypoxemia during anesthesia of wildlife. Journal of Zoo and Wildlife Medicine 43:67-76.

A131_131 Rauset, G. R., J. Kindberg, and J. E. Swenson. 2012. Modeling female brown bear kill rates on moose calves using Global Positioning Satellite data. Journal of Wildlife Management 76:1597-1606.

A132 Schregel, J., A. Kopatz, S. B. Hagen, H. Brøseth, M. E. Smith, S. Wikan, I. Wartiainen, P. E. Aspholm, O. Makarova, N. Polikarpova, M. Schneider, P. M. Knappskog, J. E. Swenson, I. Kojola, M. Ruokonen, J. Aspi, K. F. Tirronen, P. I. Danilov, and H. G. Eiken. 2012. Limited gene flow among brown bear populations in far Northern Europe? Genetic analysis of the east-west border population in the Pasvik Valley. Molecular Ecology 21:3474-3488.

A133_133 Martin, J., E. Revilla, P.-Y. Quenette, J. Naves, D. Allaine, and J. E. Swenson. 2012. Assessing brown bear habitat suitability in the Pyrenees: transferability across sites and linking scales to make the most of scarce data. Journal of Applied Ecology 49:621-631.

A134_134 Painer, J., A. Zedrosser, J. M. Arnemo, Å. Fahlman, S. Brunberg, P. Segerström, and J. E. Swenson. 2012. Effects of different doses of medetomidine and tiletamine-zolazepam on the duration of induction and immobilization in free-ranging yearling brown bears (Ursus arctos). Canadian Journal of Zoology 90:753-757.

A135_135 Ordiz, A., O.-G. Støen, S. Sæbø, J. Kindberg, M. Delibes, and J. E. Swenson. 2012. Do bears know they are being hunted? Biological Conservation 152:21–28.

A136_136 Jojola, S. M, F. Rosell, I. Warrington, J. E. Swenson & A. Zedrosser. 2012. Subadult brown bears (Ursus arctos) discriminate between unfamiliar adult male and female anal gland secretion. Mammalian Biology 77:363-368.

A137_137(a) Evans, A. V. Salén, O.-G. Støen, Å. Fahlman, S. Brunberg, K. Madslien, O. Fröbert, J. E. Swenson, and J. M. Arnemo. 2012. Capture, anesthesia, and disturbance of free-ranging brown bears (Ursus arctos) during hibernation. PLOS One 7(7): e40520.

A138_137(b) Steyaert. M. J. G., F. J. Hütter, M. Elfström, A. Zedrosser, K. Hackländer, M. H. Lê, W. M. Windisch, J. E. Swenson, and T. Isaksson. 2012. Faecal spectroscopy: a practical tool to assess diet quality in an opportunistic omnivore. Wildlife Biology 18:431-438.

A139 Bischof, R., E. B. Nilsen, H. Brøseth, P. Männil, J. Ozolinš, and J. D. C. Linnell. 2012. Implementation uncertainty when using recreational hunting to manage carnivores. Journal of Applied Ecology 49:824-832.

A140_140 Kuduk, K., W. Babik, K. Bojarska, E. Śliwińska, J. Kindberg, P. Taberlet, J. E. Swenson, & J. Radwan. 2012. Evolution of Major Histocompatibility Complex class I and class II genes in the brown bear. BMC Evolutionary Biology 12:197.

A141 Koren, L., D. Whiteside, Å. Fahlman, K. Ruckstuhl, S. Kutz, S. Checkley, M. Dumond, and K. Wynne-Edwards. 2012. Cortisol and corticosterone independence in cortisol-dominant wildlife. General and Comparative Endocrinology 177:113-119.

A142 Koren L, E. S. M. Ng, K. K. Soma, K. E. Wynne-Edwards. 2012. Sample preparation and liquid chromatography-tandem mass spectrometry for multiple steroids in mammalian and avian circulation. PLoS ONE 7(2): e32496.

A143_145 Stewart P., L. Campbell, S. Skogtvedt, K. A. Griffin, J. M. Arnemo, M. Tryland, S. Girling, M. W. Miller, M. A. Tranulis, and W. Goldmann. 2012. Genetic predictions of prion disease susceptibility in carnivore species based on variability of the prion gene coding region. PLoS ONE 7(12): e50623.

A144_130 Zedrosser, A., F. Pelletier, R. Bishof, M. Festa-Bianchet, and J. E. Swenson. 2013. Determinants of lifetime reproduction in female brown bears: early body mass, longevity, and hunting regulations. Ecology 94:231-240.

A145_139 Martin, J., B. Van Moorter, E. Revilla, P. Blanchard, S. Dray, P.-Y. Quenette, D. Allainé, and J.E. Swenson. 2013. Reciprocal modulation of internal and external factors determines individual movements. Journal of Animal Ecology 82:290-300.

A146_143 Dahle, B., K. Wallin, G. Cederlund, I.-L. Persson, L. S. Selvaag, and J. E. Swenson. 2013. Predation on adult moose Alces alces by European brown bears Ursus arctos. Wildlife Biology 19:165-169.

A147_147 Steyaert, S.M.J.G., J. Kindberg, J.E. Swenson, and A. Zedrosser. 2013. Male reproductive strategy explains spatiotemporal segregation in brown bears. Journal of Animal Ecology 82:836-845.

A148_144 Ordiz, A., O.-G. Støen, S. Sæbø, V. Sahlén, B. E. Pedersen, J. Kindberg, and Jon E. Swenson. 2013. Lasting behavioural responses of brown bears to experimental encounters with human. Journal of Applied Ecology 50:306-314.

A149 Revsbech, I.G.; H. Malte, O. Frobert, A. Evans, S. Blanc, J. Josefsson, A. Fago. 2013. Decrease in the red cell cofactor 2,3-diphosphoglycerate increases hemoglobin oxygen affinity in the hibernating brown bear Ursus arctos. American Journal of Physiology—Regulatory, Integrative, and Comparative Physiology 304: R43-R49.

A150_150 Elfström, M., O.-G. Støen, A. Zedrosser, I. Warrington, and J. E. Swenson. 2013. Gut retention time in captive Scandinavian brown bears Ursus arctos. Wildlife Biology 19:317-324.

A151_151 Friebe, A., A. Zedrosser, and J. E. Swenson. 2013. Detection of pregnancy in a hibernator based on activity data. European Journal of Wildlife Research 59:731-741.

A152 Sahdo, B., A. L. Evans, J. M. Arnemo, O. Fröbert, E. Särndahl, and S. Blanc. 2013. Body temperature during hibernation is highly correlated with a decrease in circulating innate immune cells in the brown bear (Ursus arctos): a common feature among hibernators? International Journal of Medical Sciences 10:508-514.

A153_153 Stenvinkel, P., O. Fröbert, B. Anderstam, F. Palm, M. Eriksson, A.-C. Bragfors-Helin, A. R. Qureshi, T. Larsson, A. Friebe, A. Zedrosser, J. Josefsson, M. Svensson, B. Sahdo, L. Bankir, and R. J. Johnson. 2013. Metabolic changes in summer active and anuric hibernating free-ranging brown bears (Ursus arctos) PLoS One 8(9): e7234.

A154 Jonzén, N., H. Sand, P. Wabakken, J. E. Swenson, J. Kindberg and O. Liberg, and , G. Chapron. 2013. Sharing the bounty—adjusting harvest to predator return in the Scandinavian human-wolf-bear-moose system. Ecological Modelling 265:140-148.

A155_155 López-Alfaro, C., C. T. Robbins, A. Zedrosser, and S. E. Nielsen. 2013. Energetics of hibernation and reproductive trade-offs in brown bears. Ecological Modelling 270:1-10.

A156_156 Steyaert S.M.J.G., C. Reusch, S. Brunberg, J. E. Swenson, K. Hackländer, and A. Zedrosser. 2013. Infanticide as a male reproductive strategy has a nutritive risk effect in brown bears. Biology Letters 9:20130624.

A157_157 Ordiz, A., R. Bischof, and J. E. Swenson. 2013. Saving large carnivores, but losing the apex predator? Biological Conservation 168:128-133.

A158 Stenvinkel, P., A. H. Jani, and R. J. Johnson.  2013.  Hibernating bears (Ursidae): metabolic magicians of definite interest for the nephrologist.  Kidney International 83:207–212.

A158b Norman, A. J., N. R. Street, and G. Spong. 2013. De Novo SNP discovery in the Scandinavian brown bear (Ursus arctos). PLoS ONE 8(11):e81012.

A159_158 Steyaert, S.M.J.G., J. Kindberg, J.E. Swenson, and A. Zedrosser. 2014. Litter loss triggers estrus in a nonsocial seasonal breeder. Ecology and Evolution ece3.935.

A160_138 Elfström, M., A. Zedrosser, O.-G. Støen, and J. E. Swenson. 2014. Ultimate and proximate causes behind the occurrence of bears close to settlements and its management consequences: a review. Mammal Review 44:5-18.

A161_161 Ordiz, A., J. Kindberg, S. Sæbø, J. E. Swenson, and O.-G. Støen. 2014. Brown bear circadian behavior reveals human environmental encroachment. Biological Conservation 173:1-9.

A162_162 Elfström, M, A. Zedrosser, K. Jerina, O.-G. Støen, J. Kindberg, L. Budic, M. Jonozovič, and J. E. Swenson. 2014. Does despotic behavior or food search explain the occurrence of problem brown bears in Europe? Journal of Wildlife Management 78:881-893.

A163 Nawaz, M. A., J. Martin, and J. E. Swenson. 2014. Identifying key habitats to conserve the threatened brown bear in the Himalaya. Biological Conservation 170:198-206.

A164 Odden, M., R. A. Ims, O.-G. Støen, J. E. Swenson, and H. P. Andreassen. 2014. Bears are simply voles writ large: social structure determines the mechanisms of intrinsic population regulation in mammals. Oecologia 175:1-10.

A165_165 Revsbech, I. G., X. Shen, R. Chakravarti, F. B. Jensen, B. Thiel, A. L. Evans, J. Kindberg, O. Fröbert, D. J. Stuehr, C. G. Kevil, and A. Fago. 2014. Hydrogen sulfide and nitric oxide metabolites in the blood of free-ranging brown bears and their potential roles in hibernation. Free Radical Biology & Medicine 73:349–357.

A166 Karamanlidis, A.A., S. Pllaha, L. Krambokoukis, K. Shore, and A. Zedrosser. 2014. Preliminary brown bear survey in southeastern Albania. Ursus 25:1-7.

A167 Cattet, M., G. Stenhouse, B. Macbeth, D. Janz, A. Zedrosser, J. E. Swenson, and M. Dumond. 2014. Quantifying long-term stress in brown bears with the hair cortisol concentration: a biomarker that may be confounded by rapid changes in response to capture and handling. Conservation Physiology 2: doi:10.1093/conphys/cou026.

A168_168 Friebe, A., A. L. Evans, J. M. Arnemo, S. Blanc, S. Brunberg, G. Fleissner, J. E. Swenson, and A. Zedrosser. 2014. Factors affecting date of implantation, parturition, and den entry estimated from activity and body temperature in free-ranging brown bears. PLoS ONE 9(7): e101410.

A169_169 Ozeki, L. M., Å. Fahlman, G. Stenhouse, J. M. Arnemo, N. Caulkett. 2014. Evaluation of the accuracy of different methods of monitoring body temperature in anesthetized brown bears (Ursus arctos). Journal of Zoo and Wildlife Medicine 45: 819-824.

A170_170 Græsli, A. R., Å. Fahlman, A. L. Evans, M. F. Bertelsen, J. M. Arnemo, and S. S. Nielsen. 2014. Haematological and biochemical reference intervals for free-ranging brown bears (Ursus arctos) in Sweden. BMC Veterinary Research 10:183.

A171_171 Kuduk, K. W. Babik, E. Bellemain, A. Valentini, A. Zedrosser, P. Taberlet, J. Kindberg, Jon E. Swenson, and J. Radwan. 2014. No evidence for the effect of MHC on male mating success in the brown bear. PLoS ONE 9(12): e113414.

A172_172 Elfström, M., M. L. Davey, A. Zedrosser, M. Müller, M. De Barba, O.-G. Støen, C. Miquel, P. Taberlet, K. Hackländer, and J. E. Swenson. 2014. Do Scandinavian brown bears approach settlements to obtain high-quality food? Biological Conservation 178:128–135.

A173 Sergiel, A., R. Maślak, A. Zedrosser, Ł. Paśko, D. L. Garshelis, S. Reljić, and D. Huber. 2014. Fellatio in captive brown bears: evidence of long-term effects of suckling deprivation? Zoo Biology 33:349-352.

A174_174 Jørgensen, P. G., J. Arnemo, J. E. Swenson, J. S. Jensen, S. Galatius, and O. Frøbert. 2014. Low cardiac output as physiological phenomenon in hibernating free-ranging Scandinavian brown bears (Ursus arctos) - an observational study. Cardiovascular Ultrasound 12:36.

A175 McKay, T., E. Sahlén, O.-G. Støen, J. E. Swenson, and G. B. Stenhouse. 2014. Wellsite selection by grizzly bears Ursus arctos in west-central Alberta. Wildlife Biology 5:310-319.

A176 Hodder, D. P., C. J. Johnson, R. V. Rea, and A. Zedrosser. 2014. Application of a species distribution model to identify and manage American black bear den habitat in central British Columbia, Canada. Wildlife Biology 20:238-245.

A177_177 Steyaert, S. M. J. G., J. Kindberg, K. Jerina, M. Krofel, M. Stergar, J. E. Swenson, and A. Zedrosser. 2014. Behavioral correlates of supplementary feeding of wildlife: can general conclusions be drawn? Basic and Applied Ecology 15:669–676.

A178 Fahlman, Å, J. M. Arnemo, J. Pringle, and G. Nyman. 2014. Oxygen supplementation in anesthetized brown bears (Ursus arctos)—how low can you go? Journal of Wildlife Diseases 50:574–581.

A179 Karamanlidis, A.A., A. Stojanov, M. de Gabriel Hernando, G. Ivanov, I. Kocijan, D. Melovski, T. Skrbinšek, and A. Zedrosser. 2014. Distribution and genetic status of brown bears in FYR Macedonia: implications for conservation. Acta Theriologica 59:119-128.

A180 Chapron, G., P. Kaczensky, J. D. C. Linnell, M. von Arx, D. Huber, H. Andrén, J. V. López-Bao, M. Adamec, F. Álvares, O. Anders, L. Balciauskas, V. Balys, P. Bedo, F. Bego, J. C. Blanco, U. Breitenmoser, H. Brøseth, L. Bufka, R. Bunikyte, P. Ciucci, A. Dutsov, T. Engleder, C. Fuxjäger, C. Groff, M. Heltai, K. Holmala, B. Hoxha, Y. Iliopoulos, O. Ionescu, G. Ivanov, J. Jeremić, K. Jerina, F. Knauer, I. Kojola, I. Kos, M. Krofel, J. Kubala, S. Kunovac, J. Kusak, M. Kutal, P. Mannil, R. Manz, E. Marboutin, F. Marucco, D. Melovski, K. Mersini, Y. Mertzanis, R. W. Mysłajek, S. Nowak, J. Odden, J. Ozolins, G. Palomero, M. Paunovic, J. Persson, H. Potočnik, P.-Y. Quenette, G. Rauer, I. Reinhardt, R. Rigg, A. Ryser, V. Salvatori, T. Skrbinšek, A. Skrbinšek-Majić, A. Stojanov, J. E. Swenson, A. Trajçe, E. Tzingarska-Sedefcheva, M. Váňa, R. Veeroja, M. Wölfl, S. Wölfl, F. Zimmermann, D. Zlatanova & Luigi Boitani. 2014. Successful recovery of large carnivores in Europe’s human-dominated landscapes. Science 346:1517-1519.

A181 Karamanlidis, A. A., M. Paunović, D. Ćirović, B. Karapandža, T. Skrbinšek, and A. Zedrosser. 2014. Population genetic parameters and status of brown bears in Serbia: implications for research and conservation. Ursus 25:34-43.

A182_182 Ozeki, L. M., N. Caulkett, G. Stenhouse, J. M. Arnemo, Å. Fahlman. 2015. Effect of active cooling and α₂-adrenoceptor antagonism on core temperature in anesthetized brown bears (Ursus arctos). Journal of Zoo and Wildlife Medicine 46:279-285.

A183_181 Sahlén, V., A. Ordiz, J. E Swenson, and O.-G. Støen. 2015. Behavioural differences between single Scandinavian brown bears (Ursus arctos) and females with dependent young when experimentally approached by humans. PLoS ONE 10(4): e0121576.

A184_176 Gosselin, J., A. Zedrosser, J. E. Swenson, and F. Pelletier. 2015. The relative importance of direct and indirect effects of hunting mortality on the population dynamics of brown bears. Proceedings of the Royal Society of London, B 282: 20141840.

A185_185 Frank, S., S. M. J. G. Steyaert, J. E. Swenson, I. Storch, J. Kindberg, H. Barck, A. Zedrosser. 2015. A "clearcut" case? Brown bear selection of coarse woody debris and carpenter ants on clearcuts. Forest Ecology and Management 348:164–173.

A186_186a Græsli, A. R., A. L. Evans, Å. Fahlman, M. F. Bertelsen, S. Blanc, and J. M. Arnemo. 2015. Seasonal variation in haematological and biochemical variables in free-ranging subadult brown bears (Ursus arctos) in Sweden. BMC Veterinary Research 11:301.

A187 Gilroy, J. J., A. Ordiz, and R. Bischof. 2015. Carnivore coexistence: value the wilderness. Science 347:382.

A188_179 Sahlén, V., A. Friebe, S. Sæbo, J. E. Swenson, and O.-G. Støen. 2015. Den entry behavior in Scandinavian brown bears: implications for preventing human injuries. Journal of Wildlife Management. 79:274–287.

A189_189 Paillard, L, K Jones, A. Evans, J. Berret, M. Jacquet, R. Lienhard, M. Bouzelboudjen, J. M. Arnemo, J. E. Swenson, and Maarten J. Voordouw. 2015. Serological signature of tick-borne pathogens in Scandinavian brown bears over two decades. Parasites & Vectors 8:398.

A190 Xenikoudakis, G., E. Ersmark, J.-L. Tison, L. Waits, J. Kindberg, J. E. Swenson, and L. Dalén. 2015. Consequences of a demographic bottleneck on genetic structure and variation in the Scandinavian brown bear. Molecular Biology 24:3441-3454.

A191_191 Ordiz, A., C. Milleret, J. Kindberg, J. Månsson, P. Wabakken, J. E. Swenson, and H. Sand. 2015. Wolves, people, and brown bears influence the expansion of the recolonizing wolf population in Scandinavia. Ecosphere 6(12): article 284.

A192 Stenvinkel, P. J. Kindberg, & O. Fröbert. 2015. Biomimetik – att efterlikna naturen för att förebygga sjukdom. Läkartidningen 112:586-590.

A193 Mills, J. A., C. Teplitsky, B. A. Lopez, A. Charmantier, P. H. Becker, T. R. Birkhead, P. Bize, D. T. Blumstein, C. Bonenfant, S. Boutin, A. Bushuev, E. Cam, A. Cockburn, S. D. Côté, F. Daunt, N. Dingemanse, B. Doligez, H. Drummond, R. H. M. Espie, M. Festa-Bianchet, F. Frentiu, J. W. Fitzpatrick, R. W. Furness, D. Garant, G. Gauthier, P. R. Grant, M. Griesser, L. Gustafsson, B. Hansson, M. P. Harris, F. Jiguet, P. Kjellander, E. Korpimäki, C. J. Krebs, L. Lens, J. D.C. Linnell, M. Low, A. McAdam, A. Margalida, J. Merilä, A. P. Møller, S. Nakagawa, J.-Å. Nilsson, I. C. T. Nisbet, A. van Noordwijk, D. Oro, T. Pärt, F. Pelletier, J. Potti, B. Pujol, D. Réale, R. F. Rockwell, Y. Ropert-Coudert, A. Roulin, J. S. Sedinger, Jon E. Swenson, M. E. Visser, S. Wanless, D. F. Westneat, A. Wilson, and A. Zedrosser. 2015. Archiving primary data: solutions for long-term studies. Trends in Ecology and Evolution 30:581-589.

A194 Berg von Linde, M., L. Arevström, and O. Fröbert. 2015. Insights from the den: how hibernating bears may help us understand and treat human disease. Clinical and Translational Science 8:601-605.

A195_195 Støen, O.-G., A. Ordiz, A. L. Evans, T. Laske, J. Kindberg, O. Fröbert, J. E. Swenson, and J. M. Arnemo. 2015. Physiological evidence for a human-induced landscape of fear in brown bears (Ursus arctos). Physiology and Behavior 152A:244-248.

A196 Schregel, J., Eiken, H. G., Grøndahl, F. A., Hailer, F., Aspi, J., Kojola, I., Tirronen, K., Danilov, P., Rykov, A., Poroshin, E., Janke, A., Swenson, J. E. and Hagen, S. B. 2015. Y chromosome haplotype distribution of brown bears (Ursus arctos) in Northern Europe provides insight into population history and recovery. Molecular Ecology, 24: 6041–6060.

A197_197 Stenset, N. E., P. N. Lutnæs, V. Bjarnadóttir, B. Dahle, K. H. Fossum, P. Jigsved, T. Johansen, W. Neumann, O. Opseth, O. Rønning, S. M. J. G. Steyaert, A. Zedrosser, S. Brunberg, and J. E. Swenson. 2016. Seasonal and annual variation in the diet of brown bears (Ursus arctos) in the boreal forest of southcentral Sweden. Wildlife Biology 22:107-116.

 A198_187 Bischof, R., H. Brøseth, and O. Gimenez. 2016. Wildlife in a politically divided world: insularism inflates estimates of brown bear abundance. Conservation Letters 9:122-130.

A199_199 Sommer, F., M. Ståhlman, O. Ilkayeva, J. M. Arnemo, J. Kindberg, J. Josefsson, C. Newgard, O. Fröbert, and F. Bäckhed. 2016. The gut microbiota modulates energy metabolism in the hibernating brown bear (Ursus arctos). Cell Reports 14:1-7.

A200_200. Steyaert, S. M. J. G., A. Zedrosser, M. Elfström, A. Ordiz, M. Leclerc, S. C. Frank, J. Kindberg, O.-G. Støen, S. Brunberg; and J. E. Swenson. 2016. Ecological implications from spatial patterns in human-caused brown bear mortality. Wildlife Biology 22:144-152.

A201_201. Evans, A. L., N. J. Singh, A. Friebe, J. M. Arnemo, T. G. Laske, O. Fröbert, J. E. Swenson, and S. Blanc. 2016. Drivers of hibernation in the brown bear. Frontiers in Zoology 13:7.

A202_194 Leclerc, M., J. Vander Wal, A. Zedrosser, J. E. Swenson, J. Kindberg, and F. Pelletier. 2016. Quantifying consistent individual differences in habitat selection. Oecologia 180:697-705.

A203_186b Esteruelas, N. F., N. Huber, A. L. Evans, A. Zedrosser, M. Cattet, F. Palomares, M. Angel, J. E. Swenson, J. M. Arnemo. 2016. Leukocyte coping capacity as a tool to assess capture and handling-induced stress in Scandinavian brown bears (Ursus arctos). Journal of Wildlife Diseases 52:S40-S53

A204_204 Johansson, M., O.-G. Støen, and A. Flykt. 2016. Exposure as an intervention to address human fear of bears. Human Dimensions of Wildlife 21:311-327.

A205 Rauset, G. R., H. Andrén, J. E. Swenson, G. Samelius, P. Segerström, A. Zedrosser, and J. Persson. 2016. National parks in northern Sweden as refuges for illegal killing of large carnivores. Conservation Letters 9:334–341.

A206_206 Esteruelas, N. F., J. Malmsten, C. Bröjer, G. Grandi, A. Linström, P. Brown, J. E. Swenson, A. L. Evans, J. M. Arnemo. 2016. Chewing lice Trichodectes pinguis pinguis in Scandinavian brown bears (Ursus arctos). International Journal for Parasitology: Parasites and Wildlife 5:134e138

A207 Mills, J. A., C. Teplitsky, B. A. Lopez, A. Charmantier, P. H. Becker, T. R. Birkhead, P. Bize, D. T. Blumstein, C. Bonenfant, S. Boutin, A. Bushuev, E. Cam, A. Cockburn, S. D. Côté, F. Daunt, N. Dingemanse, B. Doligez, H. Drummond, R. H. M. Espie, M. Festa-Bianchet, F. Frentiu, J. W. Fitzpatrick, R. W. Furness, D. Garant, G. Gauthier, P. R. Grant, M. Griesser, L. Gustafsson, B. Hansson, M. P. Harris, F. Jiguet, P. Kjellander, E. Korpimäki, C. J. Krebs, L. Lens, J. D.C. Linnell, M. Low, A. McAdam, A. Margalida, J. Merilä, A. P. Møller, S. Nakagawa, J.-Å. Nilsson, I. C. T. Nisbet, A. van Noordwijk, D. Oro, T. Pärt, F. Pelletier, J. Potti, B. Pujol, D. Réale, R. F. Rockwell, Y. Ropert-Coudert, A. Roulin, J. S. Sedinger, J. E. Swenson, M. E. Visser, S. Wanless, D. F. Westneat, A. Wilson, and A. Zedrosser. 2016 Solutions for archiving data in long-term studies - a reply to Whitlock et al. Trends in Ecology and Evolution 31:85–87.

A208 Penteriani, V., M. del Mar Delgado, F. Pinchera, J. Naves, A. Fernández-Gil, I. Kojola, S. Härkönen, H. Norberg, J. Frank, J. M. Fedriani, V. Sahlén, O.-G. Støen, J. E. Swenson, P. Wabakken, M. Pellegrini, S. Herrero, and J. V. López-Bao. 2016. Human behaviour may trigger large carnivore attacks in developed countries. Scientific Reports 6:20552

A209      Fernández-Gil, A., J. Naves, A. Ordiz, M. Quevedo, E. Revilla, and M. Delibes.  2016.  Conflict misleads large carnivore management and conservation: brown bears and wolves in Spain.  PLoS ONE 11(3): e0150541.

A210_210 Hertel, A., S. M. J. G Steyaert, A. Zedrosser, A. Mysterud, H. Lodberg-Holm, H. Wathne-Gelink, J. Kindberg, and J. E. Swenson.  2016.  Bears and berries: species-specific selective foraging on a patchily distributed food resource in a human-altered landscape.  Behavioral Ecology and Sociobiology 70:831-842.

A211_211 Leclerc, M., J. Van de Walle, A. Zedrosser, J. E. Swenson, and F. Pelletier. 2016. Can hunting data be used to estimate unbiased population parameters? A case study on brown bears. Biology Letters 12:20160197.

A212 Jessen, N., T. S. Nielsen, M. H. Vendelbo, R. Viggers, O.-G. Støen, A. Evans, and O. Frøbert. 2016. Pronounced expression of the lipolytic inhibitor G0/G1 Switch Gene 2 (G0S2) in adipose tissue from brown bears (Ursus arctos) prior to hibernation. Physiological Reports 4(8):e12781.

A213_213 Steyaert, S. M. J. G., M. Leclerc, F. Pelletier, J. Kindberg, S. Brunberg, J. E. Swenson, and A. Zedrosser. 2016. Human shields mediate sexual conflict in a top predator. Proceedings of the Royal Society B 283:20160906.

A214 Johansson, M., I. A. Ferreira, O.-G. Støen, J. Frank, and A. Flykt. 2016. Targeting human fear of large carnivores — Many ideas but few known effects. Biological Conservation 201:261–269

A215 Sahlén, E, S. Noell, C. S. DePerno, J. Kindberg, G. Spong, and J. P.G.M. Cromsigt. 2016. Phantoms of the forest: legacy risk effects of a regionally extinct large carnivore. Ecology and Evolution 6: 791-799.

A216 Welinder, K. G., R. Hansen, M. T. Overgaard, M. Brohus, M. Sønderkær, M. von Bergen, U. Rolle-Kampczyk, W. Otto, E. Bendixen, T. Lindahl, K. Arinell, A. L. Evans, J. E. Swenson, I. G. Revsbech, and O. Frøbert. 2016. Biochemical foundations of health and energy conservation in hibernating free-ranging subadult brown bear Ursus arctos. Journal of Biological Chemistry 291:22509-22523.

A217 Filipczyková, E., I.M.A. Heitkönig, A. Castellanos, W. Hantson, S.M.J.G. Steyaert. 2016. Marking behavior of Andean bears in an Ecuadorian cloud forest: a pilot study. Ursus 27:122-128.

A218_224 Evans, A. L., N. J. Singh, S. Blanc, A. Friebe, B. Fuchs, T. G. Laske, O. Fröbert, S. Brunberg, J. E. Swenson, and J. M. Arnemo. 2016. Physiological reactions to capture in hibernating brown bears. Conservation Physiology 4(1):cow061.

A219_223 Sivertsen, T, R., B. Åhman, S. M. J. G. Steyaert, L. Rönnegård, J. Frank, P. Segerström, O.-G. Støen, and A. Skarin. 2016. Reindeer habitat selection under the risk of brown bear predation during calving season. Ecosphere 7(11):e01583. 10.1002/ecs2.1583

A220_220 Hertel, A., A. Zedrosser, A. Mysterud, O.-G. Støen, S. M. J. G Steyaert, and J. E. Swenson. 2016. Temporal effects of hunting on foraging behaviour of an apex predator: Do bears forego foraging when risk is high? Oecologia 182:1019-1029.

A221_217 Norman, A. J., A. V. Stronen, G.-A. Fuglstad, A. Ruiz-Gonzales, J. Kindberg, N. R. Street, and G. Spong. 2017. Landscape relatedness: detecting contemporary, fine-scale spatial structure in wild populations. Landscape Ecology 32:181–194.

A222_214 Gosselin, J., M. Leclerc, A. Zedrosser, S. M. J. G. Steyaert, J. E. Swenson, and F. Pelletier. 2017. Hunting promotes sexual conflict in brown bears. Journal of Animal Ecology 86:35-42.

A223 Johansson, M., J. Frank, O.-G. Støen, and A. Flykt. 2017. An evaluation of information meetings as a tool for addressing fear of large carnivores. Society & Natural Resources 30:281-298.

A224_208 Ordiz, A., S. Sæbø, J. Kindberg, J. E. Swenson, and O.-G. Støen. 2017. Seasonality and human disturbance alter brown bear activity patterns: implications for circumpolar carnivore conservation? Animal Conservation 20: 51–60.

A225_225 Swenson, J. E., M. Schneider, A. Zedrosser, A. Söderberg, R. Franzén, and J. Kindberg. 2017. Challenges of managing a European brown bear population; lessons from Sweden, 1943-2013. Wildlife Biology 2017:wlb.00251

A226 Bischof, R., S. Steyaert, J. Kindberg. 2017. Caught in the mesh: roads and their network-scale impediment to animal movement. Ecography 40:1369–1380.

A227 De Barba M., C. Miquel, S. Lobreaux, P. Y. Quenette, J. E. Swenson, and P. Taberlet. 2017. High-throughput microsatellite genotyping in ecology: improved accuracy, efficiency, standardization and success with low-quantity and degraded DNA. Molecular Ecology Resources 17:492–507.

A228 Tallian, A., A. Ordiz, M. C. Metz, C. Milleret, C. Wikenros, D. W. Smith, D. R. Stahler, J. Kindsberg, D. R. MacNulty, P. Wabakken, J. E. Swenson, and H. Sand. 2017. Competition between apex predators? Brown bears decrease wolf kill rate on two continents. Proceedings of the Royal Society B 284:20162368.

A229_229 Leclerc, M., A. Zedrosser, and F. Pelletier. 2017. Harvesting as a potential selective pressure on behavioral traits. Journal of Applied Ecology 54:1941–1945.

A230_230 Esteruelas, N. F., M. Cattet, A. Zedrosser, G. B. Stenhouse, S. Küker, A. L. Evans, and J. M. Arnemo. 2017. Double-blinded, randomized comparison of medetomidine-tiletamine-zolazepam and dexmedetomidine-tiletamine-zolazepama anesthesia in free-ranging brown bears (Ursus arctos). PLoS ONE 12(1): e0170764.

A231 Penteriani, V., J. V. López-Bao, C. Bettega, F. Dalerum, M. del Mar Delgado, K. Jerina, I. Kojola, M. Krofel, and A. Ordiz. 2017. Consequences of brown bear viewing tourism: a review. Biological Conservation 206:169-180.

A232_232. Leclerc, M., S.C. Frank, A. Zedrosser, J.E. Swenson, and F. Pelletier. 2017. Hunting promotes spatial reorganization and sexually selected infanticide. Scientific Reports 7:45222.

A233 Redpath, S., J. C. D. Linnell, M. Festa-Bianchet, L. Boitani, N. Bunnefeld, A. Dickman, R. J. Gutiérrez, R. Irvine, M. Johansson, A. Majić, B. McMahon, S. Pooley, C. Sandström, A. Sjölander-Lindqvist, K. Skogen, J. E. Swenson, A. Trouwborst, J. Young, and E. J. Milner-Gulland. In press. Don’t forget to look down - collaborative approaches to predator conservation. Biological Reviews 92:2157-2163.

A234_234 Sergiel, A., K. A. Hobson, D. M. Janz, M. Cattet, A. Zedrosser, L. Kapronczai, C. Gryba, and N. Selva. 2017. Compatibility of preparatory procedures for the analysis of cortisol concentrations and stable isotope (δ13C, δ15N) ratios: a test on brown bear hair. Conservation Physiology 5(1):cox021.

A235 Penteriani, V., M. del Mar Delgado, F. Pinchera, J. Naves, A. Fernández-Gil, I. Kojola, S. Härkönen, H. Norberg, J. Frank, J. M. Fedriani, V. Sahlén, O.-G. Støen, J. E. Swenson, P. Wabakken, M. Pellegrini, S. Herrero, and J. V. López-Bao. 2016. Corrigendum: Human behaviour may trigger large carnivore attacks in developed countries. Scientific Reports 7:45250.

A236_236 Hertel, A. G., R. Bischof, O. Langvall, A. Mysterud, J. Kindberg, J. E. Swenson, and A. Zedrosser. In press. Berry production drives bottom-up effects on body mass and reproductive success in an omnivore. Oikos

A237_237 Frank, S. C., A. Ordiz, J. Gosselin, A. Hertel, J. Kindberg, M. Leclerc, F. Pelletier, S. M. J. G. Steyaert, O.-G. Støen, J. Van De Walle, and A. Zedrosser, and J. E. Swenson. 2017. Indirect effects of bear hunting; an important topic for managers. Ursus 28:150-181.

A238_238 Hertel, A., J. E. Swenson, and R. Bischof. 2017. A case for considering individual variation in diel activity patterns. Behavioral Ecology 28:1524–1531.

A239 Schregel, J., A. Kopatz, H. G. Eiken, J. E. Swenson, and S. B. Hagen. 2017. Sex-specific genetic analysis indicates low correlation between demographic and genetic connectivity in the Scandinavian Brown Bear (Ursus arctos). PLoS ONE 12(7): e0180701.

A240 Albrecht, J., K. A. Bartoń, N. Selva, R. S. Sommer, J. E. Swenson, and R. Bischof. 2017. How humans and climate change propelled the 12,000-year decline of the largest terrestrial carnivore. Scientific Reports 7:10399.

A241_241 Ordiz, A., O.-G. Støen, M. Delibes, and J. E. Swenson. 2017 Staying cool or staying safe in a human-dominated landscape: which is more relevant for brown bears? Oecologia 185:191–194.

A242_244 Bischof R., C. Bonenfant, I. M. Rivrud, A. Zedrosser, A. Friebe, T. Coulson, A. Mysterud, and J. E. Swenson. 2017. Regulated hunting causes life history makeover in bears. Nature Ecology and Evolution 2:116–123.

A243 Anijalg, P., S. Y. W. Ho, J. Davison, M. Keis, E. Tammeleht, K. Bobowik, I. L. Tumanov, A. P. Saveljev, E. A. Lyapunova, A. A. Vorobiev, N. I. Markov, A. P. Kryukov, I. Kojola, J. E. Swenson, S. B. Hagen, H. G. Eiken, L. Paule, and U. Saarma. In press. Large-scale migrations of brown bears in Eurasia and to North America during the Late Pleistocene. Journal of Biogeography

A244_242 Frank, S. C., M. Leclerc, F. Pelletier, F. Rosell, J. E. Swenson, R. Bischof, J. Kindberg, H. G. Eiken, S. B- Hagen, and A. Zedrosser. 2018. Sociodemographic factors modulate the spatial response of brown bears to vacancies created by hunting. Journal of Animal Ecology 87:247–258.

A245 Stenvinkel, P., J. Painer, M. Kuro-o, M. Lanaspa, W. Arnold, T. Ruf, P. G Shiels, and R. J. Johnson. In press. Novel treatment strategies for chronic kidney disease: insights from the animal kingdom. Nature Reviews.

A246. Cattet, M., G. B. Stenhouse, D. M. Janz, L Kapronczai, A.Zedrosser, A. Söderberg, J. E. Swenson, and J. Boulanger. In press. A preliminary study to determine if age class and the presence of offspring can be uncovered by concentrations of steroid hormones in the hair of brown bears. Conservation Physiology.
B) Books and book chapters

B1 Bjärvall, A., R. Franzén, M. Nordkvist and G. Åhman. 1990. Renar och rovdjur—rovdjurens effekter på rennäringen. Naturvårdsverket förlag, Solna. (Reindeer and predators—the effects of predators on the reindeer industry; in Swedish)

B2 Sandegren, F. and J. Swenson. 1997. Björnen—viltet, ekologin och människan. Svenska Jägareförbundet, Stockholm, Sweden. (The brown bear—the animal, ecology, and man; in Swedish). 70 pp.

B3 Sørensen, O. J., J. E. Swenson and T. Kvam. 1999. Status and management of the brown bear in Norway. Pages 86-89 in C. Servheen, S. Herrero and B. Peyton, compliers. Bears, status survey and conservation action plan. IUCN/SSC bear and polar bear specialist groups, IUCN, Gland, Switzerland and Cambridge, UK. 309 pp. (Invited chapter)

B4 Swenson, J. E., F. Sandegren, A. Bjärvall, R. Franzén, A. Söderberg and P. Wabakken. 1999. Status and management of the brown bear in Sweden. Pages 111-113 in C. Servheen, S. Herrero and B. Peyton, compliers. Bears, status survey and conservation action plan. IUCN/SSC bear and polar bear specialist groups, IUCN, Gland, Switzerland and Cambridge, UK. 309 pp. (Invited chapter)

B5 Swenson, J. 2000. Der Braunbär (Ursus arctos) in Eurasien. Pages 89-108 in U. Gansloßer, editor. Die Bären. Filander Verlag, Fürth. 314 pp. (Invited chapter)

B6 Swenson, J. E., N. Gerstl, B. Dahle & A. Zedrosser. 2000. Action plan for the conservation of the brown bear (Ursus arctos) in Europe. Council of Europe, Report T-PVS (2000) 24. 79 pp.

B7 Kreeger, T. J., J. M. Arnemo, and J. P. Raath. 2002. Handbook of Wildlife Chemical Immobilization. International Edition. Wildlife Pharmaceuticals Inc., Ft. Collins, Colorado, USA: 412 pp.

B8 Swenson, J. E. 2003. Implications of sexually selected infanticide for hunting of large carnivores. Pages 171-189 in M. Festa-Bianchet and M. Apollonio, eds. Animal Behavior and Wildlife Management. Island Press, Washington, D. C. 371 pp.

B9. Arnemo, J. M., B. Ranheim, A. Haga, and N. E. Søli. 2004. Sedasjon, immobilisering og anestesi av pattedyr og fugler. Sider 33e-55e i: Felleskatalog 2004-05 over preparater i veterinærmedisinen. 18. utg. Felleskatalogen AS, Oslo. [Webutgave: Fellekatalogen AS. Oslo: http://www.veterinarkatalogen.no/ (09.09.04)]

B10 Swenson, J. E. and H. Andrén. 2005. A tale of two countries: large carnivore depredations and compensation schemes in Sweden and Norway. Pages 323-339 in R. Woodroffe, S. Thirgood, and A. Rabinowitz, eds. People and Wildlife: Conflict or Co-existence? Cambridge University Press. (Invited chapter)

B11 Linnell, J. D. C., C. Promberger, L. Boitani, J. E. Swenson, U. Breitenmoser and R. Andersen. 2005. The linkage between conservation strategies for large carnivores and biodiversity: the view from the “half-full” forests of Europe. Pages 381-399 in J. C. Ray, K. H. Redford, R. S. Steneck and J. Berger . Large carnivores and the conservation of biodiversity. Island Press, Washington, D. C. 562 pp.

B12 Arnemo, J. M., B. Ranheim, A. Haga, A. J. Smith, and N. E. Søli. 2006. Sedasjon, immobilisering og anestesi av pattedyr og fugler. Sider 33e-55e i: Felleskatalog 2006-07 over preparater i veterinærmedisinen. 19. utg. Felleskatalogen AS, Oslo. [Webutgave: Fellekatalogen AS. Oslo: http://www.veterinarkatalogen.no/ (11.10.06)]

B13 Kreeger, T. J. and J. M. Arnemo. 2007. Handbook of Wildlife Chemical Immobilization. 3rd Edition. Terry J. Kreeger, Wheatland, Wyoming.

B14 Caulkett, N. A. and J. M. Arnemo. 2007. Chemical immobilization of free-ranging terrestrial mammals. Pp. 807-831 in: W. J. Tranquilli, J. C., Thurmon, and K. Grimm K, editors. Lumb & Jones’ Veterinary anesthesia and analgesia. 4th ed. Blackwell Publications, Ames.

B15 Arnemo, J. M and N. A. Caulkett. 2007. Stress. Pp. 103-109 in: G. West, D. Heard, and N. A. Caulkett, editors. Zoo animal and wildlife anesthesia and immobilization. Blackwell Publications, Ames.

B16 Arnemo, J. M., B. Ranheim, A. Haga, A. Lervik, A. J. Smith, and N. E. Søli. 2008. Sider 33e-55e i: Felleskatalog 2008-09 over preparater i veterinærmedisinen. 20. utg. Felleskatalogen AS, Oslo. [Webutgave: Fellekatalogen AS. Oslo: http://www.veterinarkatalogen.no/ (24.10.2008)]

B17 Martin, J., V. Tolon, B. van Moorter, M. Basille and C. Calenge. 2009. On the use of telemetry in habitat selection studies. Pp 37-55 in D. Barculo and J. Daniels, eds. Telemetry: research, technology and applications, Nova Science Publishers Inc, New York, USA.

B18 Arnemo, J. M., B. Ranheim, A. Haga, A. Lervik, A. J. Smith, and N. E. Søli. 2010. Sedasjon, immobilisering og anestesi av pattedyr og fugler. Sider 33e-55e i: Felleskatalog 2010-11 over preparater i veterinærmedisinen. 21. utg. Felleskatalogen AS, Oslo, 2010. [Webutgave: Fellekatalogen AS. Oslo: http://www.veterinarkatalogen.no/ (26.10.2010)]

B19 Kreeger, T. J. and J. M. Arnemo. 2012. Handbook of Wildlife Chemical Immobilization. 4th ed. Terry J. Kreeger, Sybille, Wyoming.

B20 Arnemo, J. M., B. Ranheim, A. Haga, A. Lervik, A. J. Smith, and N. E. Søli. 2012. Sedasjon, immobilisering og anestesi av pattedyr og fugler. Sider 21e-42e i: Felleskatalog 2012-13 over preparater i veterinærmedisinen. 22. utg. Felleskatalogen AS, Oslo. [Webutgave: Fellekatalogen AS. Oslo: http://www.felleskatalogen.no/medisin-vet/sedasjon-pattedyr-fugler (07.10.2012)]

B21 Arnemo, J. M., A. L. Evans, Å. Fahlman, and N. Caulkett. 2014. Field emergencies and complications. Pp. 139-147 in: G. West, D. Heard, and N. A. Caulkett, editors. Zoo animal and wildlife anesthesia and immobilization. 2nd edition. Wiley-Blackwell, Ames.

B22 Arnemo, J. M., A. Haga, A. Lervik, and N. E. Søli. 2014. Sedasjon, immobilisering og anestesi av pattedyr og fugler. Sider 21e-40e i: Felleskatalog 2014-15 over preparater i veterinærmedisinen. 23. utg. Felleskatalogen AS, Oslo. [Webutgave: Fellekatalogen AS. Oslo: http://felleskatalogen.no/medisin-vet/sedasjon-pattedyr-fugler (23.10.2014)]

B23. Swenson, J. E. 2015. Conservation research and animal activism: in search of a better balance. Pp 207-215 in L. Kemmerer (ed). Animals and the Environment: Advocacy, Activism, and the Quest for Common Ground, Routledge, London and New York.

C) Papers in nonrefereed publications

C1 Swenson, J. 1994. Sweden and Norway: historic and present status of the brown bear in Scandinavia. Intern. Bear News 3(3):5-6.

C2 Dalen, L., Johansen, T. Dahle, B. 1996. Yet another bryophyte consuming beast. Bryolog. Times 87 (2):1.

C3 Arnemo, J. M. & P. Dypsund. 1997. Kirurgisk implantation av radiosändare på vilda rovdjur. Svensk Veterinärtidning 49(1):17-18. (Surgical implantation of radio transmitters in wild carnivores. In Swedish)

C4 Arnemo, J. M. & P. Dypsund. 1997. Ny merkemetodikk på brunbjørn. Norsk Veterinærtidsskrift 109:440. (A new method for marking brown bears. In Norwegian)

C5 Sandegren, F. & J. Swenson. 1997. Det skandinaviska björnprojektet. Pp. 76-81 in Rovviltforvaltning: problemer og utfordringer. Nordisk Jägersamvirke, Rapport Nr. 4-1997. (The Scandinavian Brown Bear Research Project. In Swedish)

C6 Gossow, H. and J. Swenson. 1997. Large predator meeting in Austria. Intern. Bear News 6(3):12-13.

C7 Swenson, J. and C. Servheen. 1997. Does bear conservation without hunting produce problem bears? Intern. Bear News 6(4):11.

C8 Swenson, J. E. 1998. Coordination of large-carnivore monitoring, management, and research in Scandinavia. Pages 85-88 in C. Breitenmoser-Würster, C. Rohner, and U. Breitenmoser, editors. The re-introduction of the lynx into the Alps, Council of Europe Publishing, Environmental Encounters No. 38.

C9 Arnemo, J. M., P. Dypsund, F. Berntsen, S. J. Wedul, B. Ranheim & L. Lundstein. 1998. Bruk av implanterbare radiosendere på store rovdyr. Norsk Veterinærtidsskrift 110:799-803. (Use of implantable radio transmitters in large carnivores. In Norwegian)

C10 Andersen, R. & J. Swenson. 1999. Wildlife and nature conservation in Scandinavia with special regard to large predators. Pages 59-67 in Natura 2000 Eine Chance für den Naturschutz Europas/A chance for nature conservation in Europe. Schriftenreihe des Bundesministerium für Umwelt, Jugend und Familie, Band 14/1999. Vienna.

C11 Swenson, J. & F. Sandegren. 1999. Den svenske bjørnebestandens levedyktighet. Pages 85-90 in T. Ebenhard & M. Höggren, eds. Livskraftiga rovdjurstammar. Centrum för biologisk mångfold, Skriftserie 1. Uppsala, Sweden. (The viability of the Swedish brown bear population. In Norwegian)

C12 Swenson, J. E. and F. Sandegren. 2000. Conservation of European brown bear populations: experiences from Scandinavia. Pages 111-116 in J. F. Layna, B. Heredia, G. Palomero and I. Doadrio, editors. La conservación del oso pardo en Europa: un reto de cara al siglo XXI. Serie encuentros I. Fundación Biodiversidad, Ministerio de medio ambiente, Madrid, Spain.

C13 Dahle, B. 2000. Extensive illegal killing of brown bears in Sweden. Intern. Bear News 9(2):7.

C14 Dahle, B. 2000. Large-carnivore white paper presented in Sweden. Intern. Bear News 9(2):8.

C15 Arnemo, J. M. 2001. Reversible anaesthesia in brown bears. DDA News 2:8.

C16 Swenson, J. E. 2001. Bjørnens tilbakekomst. Det norske videnskaps-akademi, Årbok 2000: 463-470. (In Norwegian: The return of the brown bear)

C17 Arnemo, J. M., S. Brunberg, P. Ahlqvist, R. Franzén, A. Friebe, P. Segerström, A. Söderberg, and J. E. Swenson. 2001. Reversible immobilization and anesthesia of free-ranging brown bears (Ursus arctos) with medetomidine-Tiletamine-Zolazepam and Atipamezole: a review of 575 captures. Proceedings of the American Association of Zoo Veterinarians, American Association of Wildlife Veterinarians, Association of Reptilian and Amphibian Veterinarians, and National Association of Zoo and Wildlife Veterinarians Joint Conference. 2001:234-236.

C18 Zedrosser, A. & B. Dahle. 2002. Brown bear attack in central Sweden. Int. Bear News 11:9.

C19 Bjärvall, A. 2003. Stora rovdjur—samspel mellan forskning och förvaltning med björnen som exempel. Pages 13-20 in L. Terenius, Chefsredaktör. Icke-traditionella försöksdjur I forskningen. Centrala försöksdjursnämnden, Skriftserie Nr. 48. Stockholm.

C20 Swenson, J. And F. Sandegren. 2003. A summary of results from the Scandinavian brown bear research project. Pages 186-189 in P. I. Danilov and V. B. Zimin, editors. Dynamics of game animals populations in Northern Europe—Proceedings of the third international symposium. Karel’skii Nauchnyi Tsentr RAN, Redaktsionno-izdatel’skii Otdel, Petrozavodsk, Russia.

C21 Karamanlidis, A., Georgiadis, L., and Zedrosser, A. 2009. The "Southwestern Balkans Bear Register": a tool in the conservation of brown bears in the southwestern Balkans. International Bear News 18: 13-14.

C22 Swenson, J. E., A. Zedrosser, and R. Bischof. 2013. Potential evolutionary consequences of hunting brown bears. Pages 1-4 in M. Beuković, Editor-in-Chief. Modern aspects of sustainable management of game populations. The second international symposium on hunting. Faculty of Agriculture, University of Novi Sad, Novi Sad, Serbia. 268 pp.
D) Completed student theses