The scandinavian brown bear research project



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Doctorate level theses


D1.1 2003 * Bjørn Dahle, Norwegian University of Science and Technology, Trondheim. Dr. scient. (PhD) thesis: Reproductive strategies in Scandinavian brown bears.

D1.2 2004 * Jon E. Swenson, Institut für Wildbiologie und Jagdwirtschaft, Universität für Bodenkultur Wien, Vienna, Austria. Habilitation (Dr.habil.) thesis: The ecology of an increasing brown bear population: managing a successful recovery.

D1.3 2004* Alain Cercueil, Laboratoire des Techniques de l’Imagerie, de la Modélisation et de la Cognition & Laboratoire d’Ecologie Alpine, Université Joseph Fourier, Grenoble, France. Thèse d’université (PhD thesis): Contributions statistiques en génétique des populations.

D1.4 2004 * Eva Bellemain, Agricultural University of Norway, Ås (PhD) and Université Joseph Fourier, Grenoble (Docteur); joint (co-tutelle) degree program. PhD thesis/thèse d’université: Genetics of the Scandinavian brown bear: implications biology and conservation.

D1.5 2006 * Ole-Gunnar Støen, Norwegian University of Life Sciences, Ås. PhD thesis: Natal dispersal and social organization in brown bears.

D1.6 2006* Andreas Zedrosser, Norwegian University of Life Sciences, Ås and University of Natural Resources and Applied Life Sciences, Vienna. PhD thesis: Life-history strategies of brown bears.

D1.7 2006* Jonna Katajisto, University of Helsinki. Dr. scient. (Doctor of Science) thesis: Habitat use and spatial population dynamics of brown bears (Ursus arctos) in Scandinavia.

D1.8 2008* Alice Valentini, Università delgi studi della Tuscia di Viterbo (Dottorato di Recerca) and Université Joseph Fourier, Grenoble (Docteur); joint (co-tutelle) degree program. Doctoral thesis: Non-invasive diet analysis based on DNA barcoding: the Himalayan brown bears (Ursus arctos isabellinus) as a case study.

D1.9 2008* Mohammad Ali Nawaz, Norwegian University of Life Sciences, Ås. PhD thesis: Ecology, genetics and conservation of Himalayan brown bears.

D1.10 2008* Åsa Fahlman, Swedish University of Agricultural Sciences, Uppsala. PhD thesis: Advances in wildlife immobilisation and anaesthesia; clinical and physiological evaluation in selected species.

D1.11 2009* Jodie Martin, Norwegian University of Life Sciences, Ås and Université Claude Bernard – Lyon 1 / Université de Lyon, Lyon. PhD thesis: Habitat selection and movement by brown bears in multiple-use landscapes.

D1.12 2009* Richard Bischof, Norwegian University of Life Sciences, Ås. PhD thesis: Selective harvesting: short- and long-term consequences for wild populations and their management.

D1.13 2010* Andrés Ordiz, Norwegian University of Life Sciences, Ås and Universidad de León, León. PhD thesis: Reproductive patterns and human-influenced behavior in brown bears. Implications for the conservation of large carnivores.

D1.14 2010* Jonas Kindberg, Swedish University of agricultural Sciences, Umeå, PhD thesis: Monitoring and management of the Swedish brown bear (Ursus arctos) population.

D1.15 2011* Susan M. Jojola, Norwegian University of Life Sciences, Ås. PhD thesis: Social organization and olfactory communication in brown bears, Eurasian beavers, and yellow-bellied marmots.

D1.16 2012* Sam Steyaert, Norwegian University of Life Sciences, Ås and University of Natural Resources and Applied Life Sciences, Vienna. PhD thesis: The mating system of the brown bear in relation to the sexually selected infanticide theory.

D1.17 2013* Veronica Sahlén, Norwegian University of Life Sciences, Ås. PhD thesis: Encounters between brown bears and humans in Scandinavia—contributing factors, bear behavior and management perspectives.

D1.18 2013* Marcus Elfström, Norwegian University of Life Sciences, Ås. PhD thesis: Patterns and mechanisms behind the occurrence of bears close to human settlements.

D1.19 2015* Andrea Friebe, Goethe-Universität in Frankfurt am Main. PhD thesis. Winter ecology of free-ranging brown bears (Ursus arctos) in central Sweden. Dissertation.

D1.20 2015* Julia Schregel, Norwegian University of Life Sciences, Ås. PhD thesis: Genetic structure and gene flow in a continuously distributed large terrestrial carnivore – the brown bear (Ursus arctos) in Northern Europe.

D1.21 2015 Marine Plumel, Institut pluridisciplinaire Hubert Curien, Strasbourg, France. PhD thesis: Optimization of quantitative proteomics analytical strategies: application to studying metabolic adaptive responses in various organisms.

Master of Science level theses (74) (bold = accepted to a doctoral program, 20 = 27%)


D2.01 1990 Marie Dahlström, Stockholm University. Undergraduate thesis (20 points): Licensjakten på björn 1981-89—en sammanställning. (License hunting of bears, 1981-89—a summary)

D2.02 1993 Caroline Paulson, Veterinary College, Swedish University of Agricultural Sciences, Uppsala. Thesis. Immobilisering av brunbjörn. (Immobilization of brown bears)

D2.03 1995* Jens Thomas Sagør, University of Trondheim. Cand. scient. (Master of Science) thesis: Et studie av konflikten mellom bjørn og sau i perioden 1981-1993. (A study of the conflict between bears and sheep in the period 1981-1993).

D2.04 1996* Bjørn Dahle, Norwegian University of Science and Technology, Trondheim. Cand. scient. (Master of Science) thesis: Nutritional ecology of brown bears (Ursus arctos) in Scandinavia with special reference to moose (Alces alces)

D2.05 1996* Raili Ytterberg, Stockholm University. Undergraduate thesis (10 points): Do ants support the high reproductive rate in the Scandinavian brown bear (Ursus arctos) population?

D2.06 1997* Anna Jansson, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Can a high protein availablility explain the high reproductive rate in the Scandinavian brown bear (Ursus arctos)?

D2.07 1997* Thomas Johansen, Norwegian University of Science and Technology, Trondheim. Cand. scient. (Master of Science) thesis: The diet of the brown bear (Ursus arctos) in central Sweden.

D2.08 1998* Inga-Lill Persson, University of Oslo. Cand. scient. (Master of Science) thesis: Brown bear Ursus arctos predator upon adult moose in Scandinavia: a study at two levels of scale.

D2.09 1998* Ole Opseth, Norwegian University of Science and Technology, Trondheim. Cand. scient. (Master of Science) thesis: Brown bear (Ursus arctos) diet and predation on moose (Alces alces) calves in the southern tiaga zone in Sweden.

D2.10 1998* Helena Busk, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Brown bear (Ursus arctos) predation on moose (Alces alces) calves in a Swedish boreal forest.

D2.11 1999* Line Stabell, University of Oslo. Cand. scient. (Master of Science) thesis: Use of ungulates by brown brown bears Ursus arctos in Scandinavia: effects of area, season, sex, age, and individual.

D2.12 2000 Torleiv Yli Myre, Agricultural University of Norway, Ås. Cand. agric. (Master of Science) thesis: Strategies for female brown bear (Ursus arctos) to avoid infanticide: activity patterns.

D2.13 2000* Andrea Friebe, Johann-Wolfgang Goethe Universität, Frankfurt am Main, Diplomarbeit (Master of Science) thesis: Das Winterverhalten und die täglichen Wanderungen von weiblichen Braunbären (Ursus arctos) in Zentralschweden.

D2.14 2000* Håkon Hustad, Agricultural University of Norway, Ås. Cand. agric. (Master of Science) thesis: The issuing of kill permits for brown bears in response to domestic sheep depredation in Norway, 1989-99.

D2.15 2000 Beverly Ann Wilson, University of Stirling, UK. Master of Science thesis: Conservation management of the brown bear , Ursus arctos, in Scandinavia: using GIS and a habitat quality index to explain the temporal and spatial variation in sub-population core areas.

D2.16 2000 Christer Zakrisson, Swedish University of Agricultural Sciences, Umeå. Undergraduate thesis (20 points): Do brown bear (Ursus arctos) females with cubs alter their movement pattern in order to avoid infanticidal males?

D2.17 2000* Rikako Fujita, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Bait-hunting for brown bear in Sweden: temporal and spatial occurrence and potential effects on the population.

D2.18 2001 Wiebke Neumann, Universität Osnabrück, Osnabrück, Diplomarbeit (Master of Science) thesis: The brown bear Ursus arctos and berries in the Swedish boreal forest: exploiting a temporarily and spatially variable resource.

D2.19 2001 Oddmund Rønning, Agricultural University of Norway, Ås. Cand. agric. (Master of Science) thesis: Spatial and temporal supply of berries and their use by brown bears in south-central Sweden.

D2.20 2001 Valgerður Bjarnadottír, Swedish University of Agricultural Sciences, Uppsala. International Master of Science thesis: Brown bear (Ursus arctos) use of three berry species in central Sweden: do bears choose foraging plots based on the densities of berries?

D2.21 2001 Kristin Høivik Fossum, Agricultural University of Norway, Ås. Cand. scient. (Master of Science) thesis: Brown bears (Ursus arctos) and berries: use of a spatially and temporally variable resource by bears of different age and sex.

D2.22 2001 Jonna Katajisto, University of Helsinki, Helsinki. Master of Science thesis: Bed site selection of female brown bears (Ursus arctos) as a counter-strategy to avoid sexually selected infanticide by males.

D2.23 2001 * Eva Bellemain, Université Claude Bernard, Villeurbanne, France. Master of Science thesis: Etude du système d'appariement de l'ours brun de Scandinavie. (Study of the Scandinavian brown bear mating system) (In Franch and English).

D2.24 2001 Ander Larunbe Anderson, University of Stirling, UK. Master of Science thesis: Use of GIS to study brown bear, Ursus arctos, distribution in relation to habitat and traditional reindeer (Rangifer tarandus) herding in Scandinavia.

D2.25 2002 Solveig Kristoffersen, University of Oslo, Oslo. Cand. scient. (Master of Science) thesis: Restricting daily movements as a counterstrategy against sexually selected infanticide in brown bears (Ursus arctos).

D2.26 2002 Paul Antoni Nilsen, Agricultural University of Norway, Ås. Cand. scient. (Master of Science) thesis: Scandinavian brown bear (Ursus arctos L.) foraging on tempory and spatially variable berry resources in the boreal forest.

D2.27 2002 Jessica Åsbrink, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis: Parasites and metals in the Scandinavian Brown Bear (Ursus arctos). Study project from the Department of Pathology, No. 30.

D2.28 2002 Lars Plahn, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Avoidance of potentially infanticidal males by female brown bears (Ursus arctos): patterns of diel activity.

D2.29 2003 Patrik Jigsved, Umeå University, Umeå. Undergraduate thesis (30 ECTS): Utilisation of berries by brown bears in the Scandinavian boreal forest.

D2.30 2003* Knut Håkon Solberg & Ola-Mattis Drageset, Agricultural University of Norway, Ås. Cand. scient. (Master of Science) thesis: A test of methods to estimate brown bear (Ursus arctos) population size.

D2.31 2004 Insa Bauhaus, Albert-Ludwigs-Universität, Freiburg, Germany. Diplomarbeit thesis: Factors for the use of carpenter ants (Campanotus spp.) on clear-cuts by Scandinavian brown bears (Ursus arctos L.).

D2.32 2004 Eskil Nerheim, Agricultural University of Norway, Ås. Master of Science thesis: Futility of shooting brown bears Ursus arctos to stop sheep loss in Norway is confirmed.

D2.33 2004* Marcus Elfström, Umeå University, Umeå. Undergraduate thesis (30 ECTS): Denning ecology of Scandinavian brown bears.

D2.34 2004 Knut Madslien, Norwegian School of Veterinary Sciences, Oslo. Student thesis: Use of intraperitoneal radiotransmitters in yearling female brown bears; anesthetic and surgical procedures.

D2.35 2005* Ingela Jansson, Swedish University of Agricultural Sciences, Umeå. Undergraduate thesis (20 points): A pilot study of brown bear (Ursus arctos) habitat use in central Sweden using GPS.

D2.36 2005* Hanna Barck, Swedish University of Agricultural Sciences, Uppsala. Undergraduate thesis (20 points): Factors affecting foraging by brown bears Ursus arctos on carpenter ants, Camponotus herculeanus.

D2.37 2005 Claudio Signer, University of Basel, Switzerland. Master of Science thesis: Multi-level habitat analsis of female brown bears (Ursus arctos) in central Sweden.

D2.38. 2005* Therese Fosholt Moe, Norwegian University of Life Sciences, Ås. Master thesis: Diel variation in habitat selection of female Scandinavian brown bears (Ursus arctos) in relation to resting and foraging behavor.

D2.39 2006 Veronica Sahlén, University College London. Master of Science thesis: Female brown bear Ursus arctos use of an anthropogenic food source: a study from south-central Sweden

D2.40 2006* Geir Rune Rauset, Norwegian University of Life Sciences, Ås. Master thesis: Estimating individual kill rates on moose calves by brown bears based on GPS technology and GIS cluster analysis.

D2.41 2006 Inger Marie Yri, Norwegian University of Life Sciences, Ås. Master thesis: Seasonal and diel variation in road avoidance behaviour of female Scandinavian brown bears.

D2.42 2007* Bjørn Erik Pedersen, Norwegian University of Life Sciences, Ås. Master thesis: Immediate and delayed behaviour of Scandinavian female brown bears when encountered by humans on foot

D2.43 2007 Kent Ove Moren, Norwegian University of Life Sciences, Ås. Master thesis: Brunbjørns Ursus arctos predasjon på elg Alces alces: Økonomiske effekter for grunneiere i sentrale Sør-Sverige (In Norwegian: Predation by brown bears on moose: economic effects for landowners in central southern Sweden).

D2.44 2007 Jee-Yun Hyun, Seoul National University, Seoul, Republic of Korea. Masters thesis: Habitat use of female brown bears (Ursus arctos) in central Sweden (In Korean).

D2.45 2007 Maria Collett Knagenhjelm, Norwegian University of Life Sciences, Ås. Master thesisi: Bears or tears? A study of people- predator conflicts in the Bolivian Andes.

D2.46 2008* Bård Andreas Lassen, Kristian Ingdal , Telemark University College, Bø. Master of Science thesis: Chemical communication in yellow-bellied marmot (Marmota flaviventris) and brown bear (Ursus arctos).

D2.47 2008 Diala Dathe, Friedrich-Alexander Universität, Erlangen-Nürnberg, Germany. Diplomarbeit: Das Markierungsverhalten von Braunbären in süd-zentral Schweden.

D2.48 2008 Peter Marcus K. Greve, Norwegian University of Life Sciences, Ås. Master thesis: Behavior of the Scandinavian brown bear (Ursus arctos) when approached by people on foot.

D2.49 2008* Lisa Grenlund Langebro, Norwegian University of Life Sciences, Ås. Master thesis: Brown bear (Ursus arctos) bed-site selection and behaviour in relation to human settlements.

D2.50 2009* Gro Kvelprud Moen, Norwegian University of Life Sciences, Ås. Master of Science thesis: Responses by Scandinavian brown bears (Ursus arctos) to human approaches on foot.

D2.51 2009* Silje Vang, Norwegian University of Life Sciences, Ås. Master thesis: Factors affecting the ability and accuracy of dogs (Canis familiaris) to track brown bears (Ursus arctos).

D2.52 2009* Sam M. J. G. Steyaert, Wageningen University, Wageningen, Netherlands, Master of Science thesis, Habitat modelling as a predictive tool in human-wildlife conflicts: Brown bear (Ursus arctos) and free-ranging cattle in central Sweden.

D2.53 2010 Solveig Silset Berg, Norwegian University of Life Sciences, Ås. Master thesis: Small-scale habitat use by male and female brown bears (Ursus arctos) in the mating and the hyperphagy season.

D2.54 2010* Lara Budic, Norwegian University of Life Sciences, Ås. Master thesis: Brown bears in northern and southern Europe: are they seeking food or avoiding dominant bears?

D2.55 2010* Ellinor Sahlén, Swedish University of Agricultural Sciences, Umeå, Master thesis in Biology (30 points): Do grizzly bears use or avoid wellsites in west-central Alberta, Canada?

D2.56 2011* Shane Frank, University of Freiburg, Freiburg, Germany, Master of Science thesis in Forest Ecology and Management: Bears, clear-cuts and ants: predicting bear use and movement among and within clear-cuts in Sweden during summer.

D2.57 2011 David Haberkorn, Wageningen University, Wageningen, Netherlands, Master of Science thesis, Spatio-temporal analysis of brown bear (Ursus arctos) interactions in the mating season.

D2.58 2011* Cyril Milleret, Université Joseph Fourier, Grenoble, France, Master of Science thesis in Biodiversité – Écologie – Environnement. Estimating wolves (Canis lupus) and brown bear (Ursus arctos) interactions in Central Sweden. Does the emergence of brown bears affect wolf predation patterns?

D2.59 2011 Vera Menges, Edinburgh Napier University, Edinburgh. Master of Science thesis, Influence of season, sex, age and behaviour on the defecation rate of brown bears (Ursus arctos) in south-central Sweden.

D2.60 2011* Ofelia Gonzalez, Universitè de Sherbrooke, Sherbrooke, Quebec, Canada, Master thesis. Causes et consequences de la variation en tille de portée chez l’ours brun (Ursus arctos). (The determinants and effects of litter size and cub sex ratio in brown bears).

D2.61 2012* Franziska Johanna Hütter. University for Natural Resources and Applied Life Sciences, Austria, Master thesis. Fecal analysis suggests that female brown bears with dependent offspring avoid adult males during the mating season to prevent sexually selected infanticide

D2.62 2012 Erwin Pier. Wageningen University, Wageningen, Netherlands, Master of Science thesis. Impact of climate change on bear-movement regarding berry foraging; Feedback on spatio-temporal data analyses.

D2.63 2012 Jürgen Joseph, Humboldt-Universität zur Berlin. Master of Science thesis: Zur Physiologie des Infantizides bei einem nicht sozial lebenden Carnivoren, dem Braunbären (Ursus arctos) in einer frei lebenden skandinavischen Population (On the physiology of infanticide in a non-social carnivore, the brown bear (Ursus arctos) in a Scandinavian free ranging population.

D2. 64 2012* Martin Müller. University of Natural Resources and Applied Life Sciences, Vienna. Master of Science thesis: Does food matter? Diet quality of Scandinavian brown bears in relation to their sex, age, reproductive status, and distance to settlements.

D2. 65 2012 Eva Filipczyková. Wageningen University, Wageningen, Netherlands, Master of Science thesis. To feed or to leave: The effects of bait sites on the behaviour of brown bears towards humans.

D2. 66 2012. Tommy Vestøl. Norwegian University of Life Sciences, Ås. Master thesis: Baiting for brown bears (Ursus arctos) in Sweden.

D2. 67 2013. Anna Maria Petré. Swedish University of Agricultural Sciences, Umeå. Master thesis in Biology (30 points): Bears (Ursus arctos) use of managed forests in autumn; a comparison between Sweden and Canada.

D2.68 2013* Anne Randi Græsli. University of Copenhagen. Master’s thesis in Veterinary Medicine: Haematological and biochemical reference values for medetomidine-tiletamine-zolazepam immobilized free-ranging brown bears (Ursus arctos) in Scandinavia and the influence of seasons and host characteristics.

D2.69 2013* Núria Fandos Esteruelas. Universidad de Pablo Olavide, Sevilla. Master’s thesis in Biodiversity and conservation biology: Leukocyte coping capacity as a tool for assessing capture and handling-induced stress in free-ranging Scandinavian brown bears (Ursus arctos).

D2.70 2013* Christine Reusch. University for Natural Resources and Applied Life Sciences, Vienna, Austria Master thesis: Habitat segregation in brown bears: testing the predation risk hypothesis with nutritional analyses.

D2.71 2014. Lisa A. Klestil. Universität Wien. Masterarbeit: Placental scar counts and the assessment of reproductive performance in female brown bears.

D2.72 2014* Jacinthe Gosselin. Université de Sherbrooke, Sherbrooke, Québec, Canada. Grade de maître ès sciences (M.Sc.). Influence de la chasse et de l’infanticide sur la dynamique de la population de l’ours brun scandinave (Ursus arctos).

D2.73 2014 Stine Emilie Nøding Hansen. Norwegian University of Life Sciences, Faculty of Environmental Science and Technology, Department of Ecology and Natural Resource Management. Master Thesis. Behavior of Scandinavian brown bears when encountered by dogs and humans.

D2.74 2015. Nina Emilie Stenset. Norwegian University of Life Sciences, Faculty of Environmental Science and Technology, Department of Ecology and Natural Resource Management. Master Thesis. Behaviour of brown bears (Ursus arctos) when repeatedly approached by humans on foot.

D2.75 2015 Juan Romairone. Pablo Olavide University, Seville, Spain. Master of Science thesis in Biodiversity and Biology of Conservation. Habitat characteristics between different clusters of wolf (Canis lupus) activity before and after brown bear (Ursus arctos) emergence in Central Sweden.

D2.76 2015 Linda Nowak. University for Natural Resources and Applied Life Sciences, Vienna, Austria Master thesis: Reproductive performance of Scandinavian female brown bears (Ursus arctos) in relation to the use of den-type.

D2.77 2015 Marion Lukkari Arnesen. Telemark University College, Bø. Master thesis: Use of experimental bait sites by wolverines.

D2.78 2015 Henriette Wathne Gelink. Norwegian University of Life Sciences, Faculty of Environmental Science and Technology, Department of Ecology and Natural Resource Management. Master Thesis: Risky berry business? Brown bear (Ursus arctos) foraging behaviour in a landscape of fear.

D2.79 2015 Hanna Kavli Lodberg-Holm. Norwegian University of Life Sciences, Faculty of Environmental Science and Technology, Department of Ecology and Natural Resource Management. Master Thesis: When the hunter becomes the hunted: Impacts of hunting on the foraging behavior of the brown bear (Ursus arctos) in Sweden.
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Bachelor of Science level theses

D3.01 1988 Paul Klute & Tom Erik Ness, Statens skogskole, Evenstad. Undergraduate report. Älvdalsbinna - arealbruk, forflyttninger og habitatbruk 1987. (Älvdalsbinna—use of area, movements and habitat use in 1987).

D3.02 1989 * Arne Söderberg, Öster-Malma jaktvårdsskola. Undergraduate final reseach report: Mortalitet hos älgkalvar under sommaren. (Moose calf mortality in summer; In Swedish)

D3.03 1992 Mats Ola Johnsson, Klarälvdalens Folkhögskola. Undergraduate two-year school final research report. Brunbjörn. En ankät undersökning över björnförekomst i Värmlands norra delar. Samt en sammanställning av det Svensk - Norska björnprojektet. (A questionnaire study of the occurrence of bears in northern Värmland and a description of the Swedish-Norwegian Brown Bear Project).

D3.04 1995 * Andreas Norin, Klarälvdalen Folkehögskola. Undergraduate two-year school final research report. Studier av näringsval och rörelsemönster hos radiomärkta brunbjörnar (Studies of food selection and movement patterns of radio-market brown bears; In Swedish)

D3.05 1997 Paul Anzjøn & Johan Henrik Castberg, Regional College in Nord-Trøndelag, Steinkjer, Norway. Undergraduate thesis: Omtalen av bjørn (Ursus arctos) i media i 1995. (Reporting about brown bears in the Norwegian media in 1995; In Norwegian).

D3.06 1998 Pilivi Vaajakari, Omnivet. Undergraduate final report: Reproduktionen hos brunbjörnar i Sverige. (Reproduction into Brown bear in Sweden. In Swedish)

D3.07 1999 Carina Sisell, Älvdalens naturbruksgymnasium. Undergraduate final report: Björnen, människan och turismen—går de att förena? (Bears, people and tourism—is it possible to combine them? In Swedish)

D3.08 2005 Kent Ove Moren, Høgskolen i Hedmark Bachelor thesis: Brun björnens (Ursus arctos) habitatvalg:kantsoner og habitatdiversitet. (Habitat selection in brown bears (Ursus arctos): edges and habitat diversity. In Norwegian)

D3.09 2009 Elinor Sahlén, Swedish University of Agricultural Sciences, Umeå (Examensarbete i biologi, 2009:15). Undergraduate thesis (15 points, C level): Brown bear (Ursus arctos) den site concealment in relation to human activity in Scandinavia





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